Nani is the Japanese word for what, and in anime it is often delivered as a shocked response to a sudden threat or revelation. The clip became a staple of meme edits through its association with the famous Fist of the North Star exchange ending in omae wa mou shindeiru, where the opponent can only reply in disbelief. Shared widely in anime compilations, remixes and reaction videos during the late 2010s, the longer full version kept circulating because it preserves the complete timing that shorter cuts lose.
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The full Nani sound effect is the shocked Japanese 'what' reaction from anime, used in memes when something unbelievable or confusing happens.
The full Nani clip delivers the complete stretched out reaction in one go, a stunned male anime voice blurting out the Japanese word for what with rising disbelief. It starts caught off guard, then hangs on the vowel so the shock feels bigger and funnier than a short cut. That drawn out timing is why it lands so well as a punchline, instantly signalling that a character has heard something absurd, threatening or completely unexpected and needs a second to process it before anything else can happen.
